What are those holes in the cargo bed of my Nissan Frontier?

The bed of my Nissan Frontier - which usually carries the camper - has two capped holes toward the cabin. What are they for?  

Capped hole in the bed of the Nissan Frontier.
One of the holes uncapped.

No, they are not for water drainage like some people say. These holes are too high. The bed drain holes are on the cargo floor - marked in blue in the previous photo. I found that the capped holes are a factory convenience designed to run utilities to the cargo area such as wires or hoses. 

There are other holes on the sidewalls of the Frontier bed.

Holes on the sidewalls of the bed of the Nissan Frontier.
Threaded holes on the sides of the Frontier's bed.        

The upper holes are for the installation of the Utili-track racks. In these racks one can secure these attachment points - link is affiliate. 

The threaded ones near the front and the back of the sidewalls are for OEM tie downs. The thread of the tie down bolts is 8 mm - 1.25. Some advice to pass a tap with the bolt measure before installing the tie down. 

We don't use any of those tie downs to hold the truck camper, so not sure about their strength. We created our owns on the floor of the truck bed.

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